Describing a rust fungus in which the various spore forms are developed on two different and usually unrelated hosts. An example is Puccinia graminis, which causes black stem-rust of grasses and cereals and also infects barberry (Berberis). The urediospores , * teliospores , and * basidiospores develop on the grass host and the pycnospores and * aeciospores develop on barberry. Compare autoecious.
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