1. A swollen air-filled region of the thallus seen in many algae of the order Fucales, notably Fucus vesiculosus (bladder wrack). The air bladders serve to increase buoyancy and may also play a part in respiration. 2. Any of the air-filled protuberances that develop by separation of the exine layers in the pollen grains of certain gymnosperms, e.g. Pinus. They give the pollen a characteristic winged appearance and assist the wind dispersal of the pollen. See also bladder .
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